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玉山群峰線

委辦成果報告

動物

113-114年度玉山國家公園塔塔加地區臺灣水鹿族群暨植被啃磨影響監測計畫

Monitoring project for Formosan sambar population and its grazing impacts on vegetation in the Tataka Area of Yushan National Park
中文摘要
鹿科動物啃食樹木與對植被的取食行為,影響了森林的樹種組成和整體的生物多樣性。近年來臺灣水鹿(Rusa unicolor swinhoei)的族群量持續增加,除了影響高山森林生態系的正常運作,亦可能影響交通安全,因此族群量的管理將是未來重要的工作之一。本計畫的目的是於2024年3月至2025年12月在玉山國家公園的塔塔加地區,藉由架設100台自動相機監測網,估算此地區臺灣水鹿的族群量。本計劃透過計算水鹿的移動速度、辨識公鹿的個體、估算相對豐度指標、架設誘餌站等方式協助族群量的估計。另外,透過架設阻絕水鹿的圍籬,進行圍籬內外啃食壓力的比較,以了解水鹿對植被的影響程度。
結果顯示水鹿平均移動速度為20 m/min,公鹿與母鹿、亞成鹿之間的移動速度有顯著差異。水鹿偏好精鹽與混合(精鹽+蚵粉)誘餌,但不偏好氣味劑,誘餌對公鹿與母鹿的吸引效果不同,會影響相對豐度值,誘餌的使用對於族群量估算並非必要。每平方公里25台相機的密度不足以在短時間內拍到所有個體,並不適合以總量計數法估計族群量。而在捕捉標放法中,以5天為一個週期最可能符合封閉族群。塔塔加地區水鹿族群量分別為2024年6-7月166隻,2024年11-12月183隻,2025年2-3月128隻,2025年6-7月286隻,呈現明顯的季節變化,平均族群量為191隻。以公鹿最大移動距離531 m爲參考,樣區有效面積為9.61 km2,平均鹿隻密度約為20隻/ km2。水鹿與黑長尾雉(Syrmaticus mikado)的相對豐度呈顯著負相關(rs = –0.215),顯示水鹿對地面活動鳥類的影響。
水鹿阻絕圍籬於2024年12月架設完成。截至2025年10月為止,圍籬內與外的箭竹高度、箭竹覆蓋度、草本物種數量、木本小苗數量、Shannon指數與Simpson指數皆未達顯著差異。2025年7月開始發現有亞成體水鹿進入圍籬內,圍網離地高度應再降低至40 cm以下。目前塔塔加地區的臺灣水鹿密度已遠超過文獻中估算的鹿科動物對植被危害的密度閾值(5.5-15隻/km2),對於水鹿族群量的控制應及早進行。本計劃建議未來水鹿族群量估算可使用高密度自動相機(25臺/km2以上)及捕捉標放法,於6-7月水鹿角況最多樣化的時候進行,以利個體辨識。圍籬實驗則應持續記錄,以量化水鹿對植被的影響,並提供環境教育的場域。
英文摘要
Bark stripping and browsing behaviors of deer on trees and vegetation affect forest species composition and overall biodiversity. In recent years, the population of Formosan sambar deer (Rusa unicolor swinhoei) in Taiwan has continued to increase, affecting not only the normal functioning of high-mountain forest ecosystems but also potentially posing risks to traffic safety. Therefore, managing the population size has become an important future task. This project aims to estimate the population size of Formosan sambar deer in the Tataka area of Yushan National Park from March 2024 to December 2025 by deploying a network of 100 camera traps. The estimation will be assisted by calculating movement speed, identifying individual males, assessing relative abundance indices, and setting up bait stations, with four estimation sessions planned in total. In addition, exclusion fences will be established to compare browsing pressure between fenced and unfenced areas, in order to evaluate the impact of sambar deer on vegetation.
The results showed that the average movement speed of Formosan sambar deer was 20 m/min, with significant differences between males and females, as well as between males and subadults. The deer showed a preference for salt and mixed (salt + oyster shell powder) baits, but not for scent attractants. The differing attractiveness of baits to males and females may potentially influence relative abundance values, suggesting that the use of bait is not essential. A camera density of 25 units per km² was found to be insufficient to capture all individuals within a short period and is therefore unsuitable for Total count estimation of population size. In the Capture-mark-recapture analysis, a 5-day sampling period most likely met the closed-population assumption. The estimated population sizes were 166 individuals in June–July 2024, 183 in November–December 2024, 128 in February–March 2025, and 286 in June–July 2025, showing clear seasonal variation, with an average population size of 191 individuals. Based on the maximum movement distance of males (531 m), the effective sampling area was calculated as 9.61 km², resulting in an average density of approximately 20 individuals per km². The relative abundance of sambar deer was significantly negatively correlated with that of the Mikado pheasant (Syrmaticus mikado) (rs = –0.215), suggesting that sambar deer may have an impact on ground-dwelling bird species..
The deer exclusion fence was completed in December 2024. As of October 2025, no significant differences were found between the inside and outside of the fence in terms of bamboo height, bamboo cover, number of herbaceous species, number of woody seedlings, Shannon diversity index, or Simpson diversity index. Since July 2025, subadult sambar deer have been observed entering the fenced area, indicating that the bottom gap of the fence should be reduced to below 40 cm. Currently, the density of Formosan sambar deer in the Tataka area has far exceeded the threshold reported in the literature(5.5-15 individuals per km²) for deer-induced vegetation damage, suggesting that population control measures should be implemented as soon as possible. It is recommended that future population estimations of sambar deer adopt a high-density camera trapping network (over 25 cameras per km²) combined with the mark–recapture method, conducted in June–July when antler conditions are most diverse to facilitate individual identification. The exclosure experiment should be continued to quantify the impact of sambar deer on vegetation and to serve as an educational site for environmental awareness.
基本資料
  • 性質: 行政及政策類
  • 計畫編號: 1366
  • 主管機關: 內政部國家公園署玉山國家公園管理處
  • 計畫期間: 113 年 3 月至 114 年 12 月
  • 執行機關: 國立屏東科技大學
  • 年度: 114
  • 領域: 生物科學類
  • 主持人: 翁國精
  • 中文關鍵字: 水鹿、族群量估算、總量計數法、捕捉標放法、植被啃食壓力
  • 英文關鍵字:Formosan sambar deer; population estimation; Total count method; Capture -mark-recapture method; bro